Your Daily Fossil

Teleoceras

Mounted specimen on display at the Harvard Museum of Natural History

Reconstruction by Roman Uchytel

When: Miocene and Pliocene (~17.5 - 4.5 million years ago, and maybe a couple million years more!)

Where: North America

What: Teleoceras was an aquatic rhinoceros. It was a very common beast in the North American Miocene. Yes, rhinos in North America! I have been eager to share with you all the amazing diversity of North American rhinos. The discovery of a tremendous amount of rhinos, not just in terms of numbers of species but their diversity, is one of the great surprises of North American paleontological expeditions in the 19th and early 20th centuries. This continent was home to  rhinos the size of modern pigs, rhinos that could run quickly, and even aquatic rhinos! Teleoceras is one of these aquatic rhinos.

Teleoceras had very short legs for a rhino and a nubby horn. This horn is actually pretty large in the scheme of things. As much as the modern rhinos are famous for their horns the vast majority of fossil rhinos show no evidence of having a horn. We can tell this via the presence or lack of a rough surface on the nasal bones. In life Teleoceras would have probably occupied a niche very simular to the modern hippopotamus.

Plesiadapis

Mounted specimen on display at the American Museum of Natural History, NYC

Reconstruction by Jay Matternes

When: Late Paleocene to Early Eocene (~ 61 - 55 millon years ago)

Where: North America and Europe

What: Plesiadapis is a small tree-dwelling mammal that was fairly comment in the late Paleocene of North America and Europe. This ancient mammalian taxon was about the size of a house cat, and though it may look very reminiscent of a squirrel it is a member of the primate family, as part of the larger group Plesiadapiformes. The latest research has shown that Plesiadapis was actually atypical for its namesake clade; this genus tended to be much larger than the average plesiadapiform and was not as well adapted for climbing as its smaller relatives, lacking a hand specially adapted for grasping. Plesiadapis could climb trees, but it would have been an arboreal quadruped, like the living squirrels, rather than a grasping locmotion as seen in most primates today. Another features reminiscent of rodents in Plesiadapis (and this is found in most of its kin) is its enlarged front teeth and the reduction or loss of teeth between these massive incisors and the grinding cheek teeth. Plesiadapis has been reconstructed as a frugivore - meaning its diet was primarily comprised of fruit. As much of North America and Europe was covered with lush sub-tropical forests during its range, Plesiadapis would have had quite a large selection of fruits to feed on. 

The placement of Plesiadapiformes has been somewhat controversial in the past decade or so. There is uniform agreement that these animals fall somewhere near the group Euarchonta within placental mammals, but exactly where has been much debated. Euarchonta contains not only primates, but also the Scandentia (tree shrews) and Dermoptera (flying lemurs). Some early studies placed plesiadapiforms closer to the dermopterans than primates, but more recent studies tend to find this clade as either the first branches to spring off the primate lineage or just outside of Euarchonta itself, as stem taxa to all three orders. One last point to make things even more confusing! The group Plesiadapiformes? It is probably not a monophyletic (natural) group in reality. It is looking more and more like that some taxa previously grouped within Plesiadapiformes fall closer to living primates than to other taxa within  the group. 

To sum up that confusing mess, Plesiadapiformes are very important in understanding primate evolution, as at least some members of this assemblage of taxa are the first animals on the primate lineage. As this lineage includes me and you there is a lot of study focused on this group right now! Nice to see animals that are primarily paleocene taxa finally getting some attention.

Uintatherium 

Mounted specimen from American Museum of Natural History, and currently part of the traveling Extreme Mammals exhibit.

Reconstruction by Charles Knight

When: Eocene (~49 to 39 million years ago)

Where: North America

What: Uintatherium  is one of the first large mammalian herbivores. It stood about 6 feet (~1.8 meters) high at the shoulder and was roughly 13 feet (~4 meters) long. This isn’t that large for an animal today, but in the Eocene it was a giant! It lived in the lush sub-tropical forests of mid-Eocene North America, most likely eating a combination of terrestrial bushes and shrubs along with aquatic plants from lakes and marshes. Uintatherium has a nasty pair of upper canines, not what you would expect from a herbivore! It is thought that these teeth were involved in sexual display, as they appear to be much larger in males than females. Uintatherium vanishes from the fossil record in the late Eocene, at about the time the temperature of North America was falling and the vegetation was thinning out. 

Uintatherium was also one of the fossils involved in the great ‘Bone Wars’ between Cope and Marsh. It was by far the largest of the fossils to come out of the Fort Bridger fossil localities in Wyoming (this fort gives its name to a land mammal age - The Bridgerian!), and thus highly prized. Cope and Marsh both applied multiple names to specimens from this region which would later prove to all belong to the same species. The name Uintatherium wasn’t even one applied by Cope OR Marsh. Joseph Leidy named this creature in 1872, just barely edging out Marsh’s names of Dinoceras and Tinoceras. So that particular battle in the bone wars was won by someone who didn’t even have much of an interesting in fighting! 

Uintatherium is not thought to have any living descendants, it is possible that the Eocene Uintatherium was the last of its kin. However, the position of Uintatherium and its brethren (grouped as the Dinocerata) within the mammal family tree is highly uncertain. They are well accepted as placental mammals, but beyond that? It is highly debated, and in my opinion, nobody has really done a rigorous enough study to support any one position over another.  

Pachycrocuta - The Giant Hyena 

Mounted specimen from the Zhoukoudian Museum, Beijing. 

Reconstruction by Mauricio Antón

When: Pliocene to Pleistocene (~ 5 million to .5 million years ago)

Where: Europe, Asia, and Africa

What:Pachycrocutais a prehistoric member of the Hyaenidae. Today hyenas are restricted to Africa and western Asia, but their fossil record has revealed they were once much more wide spread. Pachycrocuta has been found in Africa and Asia, but most specimens have been found in Europe, with many localities in the Iberian Peninsula. The largest species was Pachycrocuta brevirostris, which stood over 3 and a half feet (~100 cm) at the shoulder and is estimated to have weighed over 400 lbs (190 kg). This makes it about the size of a modern lioness! Cave deposits in both Spain and China have revealed multiple almost complete skeletons, suggesting that these animals lived in packs and utilized these caves as their dens. 

As Pachycrocuta is even more heavyset and adapted for bone crunching than the living bone-crunching hyenas, it has been suggested that this fossil form was even more dependent on scavenging kills than living species. But there really is not much evidence to pack this up other than thought-experiments. As it appears that some large cat species were displaced when Pachycrocuta moved into their ranges, it is more likely it was a direct hunter that would take advantage of pre-killed remains when it could drive away other predators. Like 99% of carnivores today. The predator/scavenger divide is really not a fast or hard line at all. Even more evidence of hunting comes from the remains of interactions between Pachycrocuta and Homo errectus. These two species overlapped and bones of our poor relative have been found in Pachycrocuta dens in China! 

Palaeolagus

When: Late Eocene to Mid Oligocene (~38 to 27 million years ago)

Where: North America

What: Palaeolagus is a fossil lagomorph. Lagomorpha is an order of mammals, that contains rabbits, hares, and pikas. Within the bunny-order rabbits and hares are more closely related to either other than either is to the pikas. If you are not familiar with pikas go check out some pictures! They are really cute little guys that resemble guinea pigs more than they do rabbits, but they are most assuredly lagomorphs. Palaeolagus falls outside all living lagomorphs in their evolutionary lineage. It can be thought of as representative of the common ancestor of all living lagomorphs.  

Palaeolagus lived in North America in the late Eocene, after the dense forests had left and the grasslands of the plains started to expand. This 10 inch (~25 cm) long herbivore spread throughout the continent during the Oligocene as the grasslands grew. Palaeolagus could not hop, its hind legs show none of the features that make a hopping locomotion style possible in living rabbits.This ancient bunny is known from a large amount of fossil specimens, some of which are almost complete skeletons, but most are fragmentary pieces of bone or teeth. Most of these Palaeolagus specimens likely met their end as the lunch of one of the many predators roaming the grass lands of prehistoric North America. 

Procoptodon - The giant short faced kangaroo 

Mounted skeleton on display at Victoria Fossil Cave, Naracoorte Caves National Park, South Australia

Reconstruction by Peter Trusler. 

When: Pleistocene (~ 2 million to 15,000 years ago)

Where: Throughout Australia

What: Procoptodon is a giant fossil kangaroo. Exactly how ‘giant’ it is has been a bit exaggerated, heights of up to 10 feet (~3 meters) have been reported, but this would have been its maximum height when it reared up fully on its hind legs, with its arms reaching up for high branches. Procoptodon was capable of this posture, but (like living kangaroos) it did not stand fully upright most of the time. In its normal feeding (and most everything else) poster it would have stood about 6.5 feet (~ 2 meters) tall; about the same height as the largest of the modern red kangaroos. Procoptodon was not the same size as these animals though, it was much more massive and would have been over twice the weight of a red kangaroo of equivalent height. 

Procoptodon was very well adapted for the semiarid conditions that characterized much of Australia during the Pleistocene, but fossil remains have also been found in the more hospitable regions of prehistorical Australia.  The marsupials of Australia are well known for their convergence evolution upon forms from other continents (such as the tasmanian tiger and the marsupial mole), but the kangaroo does not look like any placental mammal known. However, in terms of its lifestyle, the ecological niche that it inhabits, the group is convergent upon hoofed animals, such as deers!  Procoptodon overlapped with human habitation of Australia, and it is thought some Aboriginal folktales are about this massive kangaroo. 

Procoptodon is a member of the group Sthenurinae - the shortfaced kangaroos. As you probably guessed these kangaroos had much shorter snouts than the modern species of kangaroos. This group is completely extinct. It is one of the subgroups of the Macropodidae, the clade of marsupials that contains all kangaroos and wallabies, as well as a few other  groups. It has been proposed that within the Macropodidae the closest living relative of Procoptodon is the Banded hare-wallaby, though this is not universally accepted. 

In the prehistoric outback Procoptodon would have co-exsited with the largest marsupial of all time Diprodoton and was a hunted by the marsupial lion Thylacoleo. And the second link you can see this marsupial predator hunting a close relative of Procoptodon!  

Ambulocetus

Reconstructions by Carl Buell 

When: Eocene (~50 to 48 million years ago)

Where: Pakistan 

What: Ambulocetus is fossil whale relative. This beast was about 10 feet (~3 meters) long, and not very agile in either the land or the water. It was capable of movement on land, but it would have been rather slow and lumbering, as its forelimbs were shortened compared to its fully terrestrial ancestors. In the water it would have been capable of swimming with some speed, but it would not have been able to make quick turns as it chased its prey. Therefore, it has been reconstructed as an ambush-style predator, in the same niche as the modern crocodile. It would have laid in wait in the water, with its relatively dorsal eyes and nose peeking above the sufrace, able to see and smell approaching prey. Once a prey animal got close enough, Ambulocetus would launch itself from the water and try to catch the animal in its powerful jaws, such as is shown above. I think it is some form of basal horse that is trying to avoid the snapping jaws of Ambulocetus. This ambush style strategy could have also worked with aquatic prey, such as schools of fish. Ambush predation is seen in some species of whales today, Orcas (the killer whales) have been recorded ambushing seals on ice flows. 

Ambulocetus lived on the edge of the Tethys Sea (a body of water between India and Asia) in what is now Pakistan. At the time this region was one of many islands off the shore of the island continent of India, which had not yet collided with Asia (this would not happen for tens of millions of years). This warm seaway was full of mammals starting to return to the seas, including other lineages of whale relatives. In the cetacean family tree, Ambulocetus falls between Indohyus and modern whales; it was carnivorous - as all modern whales are-, and far more adapted for aquatic locomotion than Indohyus was, with shortened legs and a much more powerful tail. 

Indricotherium - The largest terrestrial mammal

Skull on display at the American Museum of History of Natural History, New York City

Reconstruction was part of the traveling Extreme Mammals exhibit, photo from when it was at the AMNH. 

When: Eocene and Oligocene (~ 34 to 23 million years ago)

Where: Asia and Eastern Europe 

What: Indricotherium is the largest terrestrial mammal known. It is a member of the rhinoceros family. Some material was found earlier in the 20th century, but the first fairly complete skull and skeletal elements were found by Roy Chapman Andrews in 1922 while on an expedition for the American Museum of Natural History in Mongolia. Indricotherium would have stood about 16.5 feet (~5 meters) tall at the shoulder and is estimated to have weighed in excess of 20 tons. There were sauropod dinosaurs that were smaller than Indricotherium! This giant was a herbivore and filled a simular niche to the sauropods and modern giraffes, so much so that it is sometimes referred to as the ‘giraffe rhinoceros’.  It stripped tall trees bare of leaves using its large front teeth and mobile lips. 

In the family tree of mammals Indricotherium is in the order Perrisodactyla (horses, rhinos, and tapirs). Within this group it in the rhinoceros clade. While rhinos today all look pretty much the same their fossil record shows this group used to be extremely diverse, with two completely extinct major sub groupings. Indricotherium is in the group Hyracodontidae (the running rhinos). This lineage divered from that leading to modern rhinos over 55 million years ago. Not all rhinos in this group were giant sized, the very first ones were no larger than wolves! All members of this group lacked horns like Indricotherium. 

There is a bit of controversy and confusion surrounding the topic of what genus name to apply to this animal. Indricotherium has been proposed by some workers to be synonymous with Paraceratherium and Baluchitherium, but this is not universally accepted. I have used the name Indricotherium for this entry as both examples shown above are based upon material that has held the name Indricotherium. 

Leptictidium

When: Eocene (~48-40 million years ago)

Where: Found at the Messel fossil site in Germany

What: Leptictidium is one of the more common mammals found in the Messel fossil pit in Germany. The adults ranged from about two (~60 cm) to three (~90 cm) feet in length, with most of this length being in the long tail. Leptictidium had extremely short forelimbs relative to the length of its legs, and has been reconstructed as the first bipedal mammal. There is debate as to its precise mode of locomotion, with some researchers proposing that the animal was a fast runner and others suggesting it was saltatorial (hopping). More recent studies have supported a hopping and leaping mode of locomotion. Thanks to the extraordinary preservation of fossils from Messel, we know the tail was bald for much of its length, that Leptictidium had a short ‘trunk’, like the elephant shrews of the modern day, and that this animal ate insects and small vertebrates. Contemporaries of Leptictidium include the tiny horse Propalaeotheirum and the predatory giant flightless bird Gastornis. 

Leptictida is the larger clade that includes Leptictidium and its kin. The first members of this group appear in the latest Cretaceous of western North America and the order quickly spreads throughout the northern continents, lasting until the early Oligocene about 30 million years ago, when the forests worldwide started to give way to grasslands. Previously leptictids were thought to be related to either the living lipotyphla (hedgehogs, shrews, and moles) or elephant shrews, but recent studies of the relationships of mammals have placed them outside of placental mammals entirely, making them stem eutherians and not members of Placentalia. 

Deinogalerix

Mounted specimen from the National Museum of Natural History in Leiden, the Netherlands. 

Reconstruction by Mauricio Anton with a common hedgehog for scale. 

When: Miocene (~11 - 5 million years ago)

Where: One island that now is part of Italy

What:Deinogalerix is a comparatively giant relative of the hedgehog. It lived on what is now the Gargano peninsula in Italy, but during the Miocene this region was a separate island. Much of Italy during this time period was a series of isolate islands, owing to the higher water level. Deinogalerix was about five times the size of a common hedgehog, more the size of a small fox. However, with a skull about 1/3rd the total length of its whole body, it was proportioned very differently. An eight inch (~20 cm) skull on a 24 inch (~60cm) body isn’t too out of proportion for many of the Lipotyphla (the order that includes hedgehogs, shrews, moles, and solenodons), and it appears Deinogalerix saw no reason to shrink down its head just because of its growth spurt.  I have called this animal a hedgehog, and it is in that grouping, but it did not look much at all like the little spiny animal shown above. Within the hedgehog family, its closet relatives are not true hedgehogs, but rather the gymnures or ‘moon-rats’. These animals have not developed spines as protection and are covered with a coat of long course hairs.